FROLOV, Andrey V.; AKHMETOVA, Lilia A. Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae). European Journal of Taxonomy, [S. l.], n. 241, 2016. DOI: 10.5852/ejt.2016.241. Disponível em: https://ejt.sandbox.publicknowledgeproject.org/index.php/ejt/article/view/365. Acesso em: 25 apr. 2026.