Genner, Martin J., et al. “Revision of the African Cichlid Fish Genus Ctenochromis (Teleostei, Cichliformes), Including a Description of the New Genus Shuja from Lake Tanganyika and the New Species Ctenochromis Scatebra from Northern Tanzania”. European Journal of Taxonomy, vol. 819, no. 1, May 2022, pp. 23-54, doi:10.5852/ejt.2022.819.1775.