Kolesnichenko, Kirill A., and Anatoly A. Kotlobay. “Review of the Fritillary Species Systematically Close to Melitaea Lutko Evans, 1932 (Lepidoptera: Nymphalidae) With Analysis of Their Geographic Distribution and Interrelations With Host Plants”. European Journal of Taxonomy 830, no. 1 (July 15, 2022): 1–60. Accessed April 24, 2026. https://ejt.sandbox.publicknowledgeproject.org/index.php/ejt/article/view/1865.