Hughes, Mark, Deden Girmansyah, and Wisnu Handoyo Ardi. “Further Discoveries in the Ever-Expanding Genus Begonia (Begoniaceae): Fifteen New Species from Sumatra”. European Journal of Taxonomy, no. 167 (December 21, 2015). Accessed April 28, 2026. https://ejt.sandbox.publicknowledgeproject.org/index.php/ejt/article/view/289.